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Camcore Book

Conservation and Testing of Tropical Subtropical Forest Tree Species by the camcore cooperative (2000)

This book describes 11 different pines in Central America and Mexico that Camcore has worked with over the years. It has a number of color photos and tables about collection sites as well as early results of many Camcore trials.  The book can be obtained by contacting the Camcore office. A nominal fee will be charged for shipping. Email us at to request a copy of the book.

Academic Publications [PDF Files]


Acosta J.J. , Fahrenkrog A.M., Neves, L.G., Resende, M.F.R., Dervinis, C., Holliday, J.A.. Kirst, M.  2018. Exome re-sequencing reveals parallel evolutionary history, genomic diversity, and targets of selection in the conifers Pinus taeda and Pinus elliottii.  Genome Biology and Evolution. Submitted.

Hodge, G.R., Acosta, J.J., Mansfield, S.D., Woodbridge, W.C.,  Unda, F. 2018. Global NIR Models to Predict Wood Chemical Properties of Eucalyptus. Journal of Near Infrared Spectroscopy.

Lapham, M., C.F. Ford, A.E. Mayfield III, R.M. Jetton, S.T. Brantley, D.R. Zietlow, C. Brown, and J.R. Rhea. 2018.  Shade and hemlock woolly adelgid infestation increase eastern hemlock foliar nutrient concentration. Forest Science.

Lopez, J.L., R.C. Abt, W.S. Dvorak, G.R. Hodge, and R. Phillips. 2018. Tree breeding model to assess financial performance of pine hybrids and pure species: deterministic and stochastic approaches for South Africa. New Forests.

Naidoo, S., Christie, N., Acosta J.J., Mphahlele, M., Payn, K., Myburg, A., Külheim, C. 2018. Terpenes associated with resistance against the gall wasp, Leptocybe invasa, in Eucalyptus grandis. Plant, Cell and Environment.

Whittier, W.A. 2018. Nutrient Disorder Foliar Symptoms, Foliar Nutrient Levels and Predictive Near-infrared Spectroscopy Nutrient Models of Teak (Tectona grandis L.f.).  NCSU Master of Science Forestry Thesis.


Brantley, S.T., A.E. Mayfield III, R.M. Jetton, C.F. Miniat, D.R. Zietlow, C. Brown, and J.R. Rhea. 2017. Elevated light levels reduce hemlock woolly adelgid infestation and improve carbon balance of infested eastern hemlock seedlings. Forest Ecology and Management.

Bucholz, E., J. Frampton, R. Jetton, D. Tilotta, and L. Lucia. 2017. Effect of different headspace concentrations of bornyl acetate on fecundity of green peach aphid and balsam woolly adelgid. Scandinavian Journal of Forest Research.

de Waal, L., R.G. Mitchell, G.R. Hodge and P.W. Chirwa. 2017. The use of field and artificial freezing studies to assess frost tolerance in natural populations of Pinus oocarpa.  Southern Forests: a Journal of Forest Science.

Hansen, O.K., S. Changtragoon, B. Ponoy, J. Lopez, J. Richard, and E.D. Kjær. 2017. Worldwide translocation of teak – origin of landraces and present genetic base. Tree Genetics and Genomes.

Hastings, J.M., K.M. Potter, F.H. Koch, M. Megalos, and R.M. Jetton. 2017. Prioritizing conservation seed banking locations for imperiled hemlock species using multi-attribute frontier mapping. New Forests.

Hongwane, Phillip, G. Mitchell, A. Kanzler, S. Verryn, J. Lopez, and P. Chirwa. 2017. Alternative pine hybrids and species to Pinus patula and P. radiata in South Africa and Swaziland, Southern Forests. A Journal of Forest Science.

Lstiburek, M, Y.A. El-Kassaby, T. Skroppa, G.R. Hodge, J.H. Sonstebo, A. Steffenrem. 2017. Dynamic gene-resource landscape management of Norway spruce: combining utilization and conservation. Frontiers in Plant Science.

Moraes, B.F.X.; Santos, R.F.,;Lima B.M.; Aguiar, A.M; Missiaggia, A.A.; Dias, D.; Rezende, G.D.; Gonçalves, F.M.; Acosta, J.J.; Kirst, M.; Resende, M.F.R.; Munoz, P.R. 2017. Comparing Sequence Capture and SNP-array genotyping methods for development of genomic selection prediction models. Molecular Breeding. Submitted.

Potter, K.M., A.R. Campbell, S.A. Josserand, C.D. Nelson, and R.M. Jetton. 2017. Population isolation results in unexpectedly high differentiation in Carolina hemlock (Tsuga caroliniana), an imperiled Southern Appalachian endemic conifer. Tree Genetics and Genomes.

Potter, K.M., R.M. Jetton, A. Bower, D.F. Jacobs, G. Man, V.D. Hipkins, and M. Westwood. 2017. Banking on the future: progress, challenges and opportunities for the genetic conservation of forest trees. New Forests.


Hastings, J.M. 2016. Using climate and genetic diversity data to prioritize conservation seed banking for imperiled hemlock species. M.S. Thesis, NC State University, Raleigh, NC. 73 p.


Hodge, G.R. and W.S. Dvorak. 2015. Provenance variation and within-provenance genetic parameters in Eucalyptus urophylla across 125 sites in Brazil, Colombia, Mexico, South Africa, and Venezuela. Tree Genetics and Genomes.

Jetton, R.M., Crane, B.S., Whittier, W.A. and Dvorak, W.S. 2015. Genetic Resource Conservation of Table Mountain Pine (Pinus pungens) in the Central and Southern Appalachian Mountains. Tree Planters’ Notes.

Lopez, J.L . 2015. Economic potential of pine hybrids: a case study for South Africa. PhD Dissertation, NC State University, Raleigh, NC.

Lstiburek, M, G.R. Hodge, and P. Lachout. 2015. Uncovering genetic information from commercial forest plantations: making up for lost time using “Breeding without Breeding”. Tree Genetics and Genomes.

Mayfield III, A.E., Reynolds, B.C., Coots, C.I., Havill, N.P., Brownie, C., Tait, A.R., and Hanula, J.L.,. Joséph, S.V., Galloway, A.B. 2014. Establishment, hybridization and impact of Laricobius predators on insecticide-treated hemlocks: Exploring integrated management of the hemlock woolly adelgid. Forest Ecology and Management. 

Powers, Z., A. Mayfield, J. Frampton, and R. Jetton. 2015. Comparison of suspended branch and direct infestation techniques for artificially infesting hemlock seedlings with the hemlock woolly adelgid for resistance screening. Forests.


Acosta, J.J., 2014. Genetic diversity and genome-wide selection studies in pines and Eucalyptus. Ph.D. dissertation, School of Forest Resources and Conservation, University of Florida. Gainesvile, FL.

Campbell, A.R. 2014. Assessing Carolina hemlock (Tsuga caroliniana Engelm.) population genetic structure anddiversity in the southern Appalachians using microsatellite markers. M.S. Thesis, NC State University, Raleigh, NC. 73 p.

Gapare, W.J. 2014. Merging applied gene conservation activities with advanced generation breeding initiatives: a case study of Pinus radiata D. Don. New Forests.

Hodge, G.R, and W.S. Dvorak. 2014. Breeding southern US and Mexican pines for increased value in a changing world. New Forests.

Jetton, R.M. 2014. Collecting seeds and managing insects: my career as a forest conservation biologist. CCES Highlights Magazine.

Jetton, R.M., Mayfied III, A.E., Powers, Z.L., 2014. Development of a Rain Down Technique to Artificially Infest Hemlocks with the Hemlock Woolly Adelgid (Hemiptera: Adelgidae). Journal of Insect Science.

Jetton, R.M., Robison, D.J., 2014. Effects of Artificial Defoliation on Growth and Biomass Accumulation in Short- Rotation Sweetgum (Liquidambar styraciflua L.) in North Carolina, USA.. Journal of Insect Science.

Jetton, R.M., Whittier, W.A., and Dvorak, W.S., 2014. Evaluation of stratification treatments for germinating seeds of Tsuga canadensis and Tsuga caroliniana for ex situ gene conservation. Southeastern Naturalist.

Kanzler, A., A.Nel, and C. Ford. 2014. Development and commercialisation of the Pinus patula × P. tecunumanii hybrid in response to the threat of Fusarium circinatum. New Forests.

Nel, A., G.R. Hodge, K.E. Mongwaketsi, and A. Kanzler. 2014. Genetic parameters for Fusarium circinatum tolerance within open-pollinated families of Pinus patula tested at screening facilities in South Africa and the USA. Southern Forests.

Oten, K.L.F., Merkle, S.A., Jetton, R.M., Smith, B.C., Talley, M.E., and Hain, F.P. 2014. Understanding and developing resistance in hemlocks to the hemlock woolly adelgid. Southeastern Naturalist.

Powers, Z.L. 2014. Evaluating techniques for artificially infesting hemlocks (Tsuga spp.) with the hemlock woolly adelgid (Adelges tsugae) for genotype resistance screening. M.S. Thesis, NC State University, Raleigh, NC 151 p.


Amborella Genome Project (Acosta, J.J.), 2013. The Amborella genome and the evolution of flowering plants. Science.

Dvorak, W.S., Nel, A., Espinoza, J.A., 2013. Evidence of low-levels of natural introgression between Pinus jaliscana and P. oocarpa in an open-pollinated progeny trial using near-infrared spectroscopy. Forest Ecology and Managment.

Jetton, R.M., 2013. Saving hemlocks, one seed at a time. Chinquapin.

Jetton, R.M., Whittier, W.A., Dvorak, W.S., Rhea, J., 2013. Conserved Ex Situ Genetic Resources of Eastern and Carolina Hemlock: Eastern North American Conifers Threatened by the Hemlock Woolly Adelgid. Tree Planters’ Notes.

Leibing, C., Singer, J., van Zonneveld, M., Jarvis, A., Dvorak, W.S., 2013. Selection of Provenances to Adapt Tropical Pine Forestry to Climate Change on the Basis of Climate Analogs. Forests.


Dvorak, W.S., 2012. The strategic importance of applied tree conservation programs to the forest industry in South Africa. Southern Forests.

Dvorak, W.S., 2012. Water use in plantations of eucalyptus and pines: a discussion paper from a tree breeding perspective. International Forestry Review.

Espinoza, J.A. Allen, H.L., McKeand, S.E., Dougherty. P.M., 2012. Stem sinuosity in Loblolly pine and nitrogen and calcium additions. Forest Ecology and Management.

Espinoza, J.A., Hodge, G.R., Dvorak, W.S., 2012. The potential use of near infrared spectroscopy to discriminate between different pine species and their hybrids. Journal of Near Infrared Spectroscopy.

Hodge, G.R., Dvorak, W.S., 2012. Growth potential and genetic parameters of four Mesoamerican pines planted in the Southern Hemisphere. Southern Forests.

Hodge, G.R., Dvorak, W.S., Tighe, M.E., 2012. Comparisons between laboratory and field results of frost tolerance of pines from the southern USA and Mesoamerica planted as exotics. Southern Forests.

Mitchell, R.G., Wingfield, M.J., Hodge, G.R., Steenkamp, E. Coutinho, T.A., 2012. Selection of Pinus spp. in South Africa for tolerance to infection by the pitch canker fungus. New Forests.

Mitchell, R.G., Wingfield, M.J., Hodge, G.R., Steenkamp, E. Coutinho, T.A., 2012. Susceptibility of provenances and families of Pinus maximinoi and Pinus tecunumanii to frost in South Africa. New Forests.

Mitchell, R.G., Wingfield, M.J., Hodge, G.R., Steenkamp, E. Coutinho, T.A., 2012. The tolerance of Pinus patula x Pinus tecunumanii, and other pine hybrids to Fusarium circinatum in greenhouse trials. New Forests.

Potter, K.M., Jetton, R.M., Dvorak, W.S., Hipkins, V.D., Rhea, J.R., Whittier. W.A., 2012. Widespread inbreeding and unexpected geographic patterns of genetic variation in Eastern hemlock (Tsuga canadensis), an imperiled North American conifer.Conservation Genetics.


Jetton, R.M., Monahan, J.F., Hain, F.P., 2011. Laboratory studies of feeding and oviposition preference, developmental performance, and survival of the predator Sasajiscymnus tsugae on diets of Adelges tsugae and Adelges piceae. Journal of Insect Science.

Wee, K.S., ChunHong, L., Dvorak, W.S., Han, Y., 2011. Genetic diversity in natural populations of Gmelina arborea: implications for breeding and conservation. New Forests.


Hodge, G.R., Woodbridge, G.R., 2010. Global near infrared models to predict lignin and cellulose content of pine wood. Journal of Near Infrared Spectroscopy.

Jetton, R.M., Rhea, J.R., 2010. Framework for genetic conservation of Eastern (Tsuga canadensis) and Carolina (T. caroliniana) hemlock. Proceedings of the Fifth Symposium on Hemlock Woolly Adelgid in the Eastern United States.

Jetton, R.M., Whittier, W.A., Dvorak, W.S., Rhea, J.R., 2010. Status of Gene Conservation for Eastern and Carolina hemlock in the Eastern United States.Proceedings of the Fifth Symposium on Hemlock Woolly Adelgid in the Eastern Untied States.

Potter, K.M., Jetton, R.M., Dvorak, W.S., Hipkins, V.D., Rhea, J.R., Whittier, W.A., 2010. Of microsatellites, Hemlock Woolly Adelgid, and climate change: Assessing genetic threats to it, across the range of Eastern hemlock. Proceedings of the Fifth Symposium on Hemlock Woolly Adelgid in the Eastern United States.

Potter, K.M., Jetton, R.M., Dvorak, W.S., Frampton, J., Rhea, J.R., 2010. Ex situ seed collections represents genetic variation in natural stands of Carolina hemlock.Proceedings of the Fifth Symposium on Hemlock Woolly Adelgid in the Eastern United States.

Schimleck, L.R. Hodge, G.R., Woodbridge, W.C., 2010. Toward global calibrations for estimating the wood properties of tropical, sub-tropical and temperate pine species.Journal of Near Infrared Spectroscopy.


Dvorak. W.S., 2009. How private industry has met the challenge of conservation of forest genetic resources of tropical and subtropical tree species. International Technical Association of Tropical Timber.

Dvorak, W.S., Potter, K.M., Hipkins, V.D., Hodge, G.R., 2009. Genetic Diversity and Gene Exchange in Pinus oocarpa, a Mesoamerican Pine With Resistance to the Pitch Canker Fungus (Fusarium circinatum). International Journal of Plant Sciences.

Jetton, R.M., Hastings, F.L., Hain, F.P., 2009. Arthropod Diversity Associated with Old and Secondary Growth Eastern hemlock (Tsuga canadensis) in the Great Smoky Mountains National Park. National Park Journal of the North Carolina Academy of Sciences.

Jetton, R.M., Dvorak, W.S., Potter, K.M., Whittier, W.A., Rhea, J.R., 2009. Genetics and conservation of hemlock species threatened by the hemlock woolly adelgid.Proceedings of the 30thSouthern Forest Tree Improvement Conference, 2009.

Leibing, C., Van Zonneveld, M., Jarvis, A., Dvorak, W.S., 2009. Adaptation of tropical and subtropical pine plantation forestry to climate change: Realignment of Pinus patulaand Pinus tecunumanii genotypes to 2020 planting site climates. Scandinavian Journal of Forest Research.

Lopez, J.L., De La Torre, R., Cubbage, F., 2009. Effect of land prices, transportation costs, and site productivity on timber investment returns for pine plantations in Colombia. New Forests.

Lopez, J.L., Gomide, J., Phillips, R., 2009. Influence of Eucalyptus wood properties on the financial performance of a modeled Brazilian pulp mill. O Papel.

Van Zonneveld, M., Jarvis, A., Dvorak, W.S., Lema, G., Leibing, C., 2009. Climate change impact predictions on Pinus patula and Pinus tecunumnaii populations in Mexico and Central America. Forest Ecology and Management.

Boletín de Noticias Camcore para México y Centroamérica

2011 – Julio (July)

2011 – Mayo (May) – Teak and Gmelina Newsletter Number 1